1. Acadia Wildlife Services, PO Box 56, South Freeport, Maine 04078, USA
2. Disease Investigations, San Diego Zoo Global, 15600 San Pasqual Valley Rd., Escondido, California 92027, USA
3. Animal Health, San Diego Zoo Global, 15500 San Pasqual Valley Rd., Escondido, California 92027, USA
4. Desert Tortoise Recovery Office, US Fish and Wildlife Service, 1340 Financial Blvd., Suite 234, Reno, Nevada 89502, USA
5. Western Ecological Research Center, US Geological Survey, 160 N Stephanie St., Henderson, Nevada 89074, USA
6. Department of Geography, University of Nevada, Reno, 1664 N Virginia St., Reno, Nevada 89557, USA