1. Southeastern Cooperative Wildlife Disease Study, College of Veterinary Medicine, University of Georgia, 589 D. W. Brooks Drive, Athens, Georgia 30602, USA
2. Department of Veterinary and Biomedical Sciences, Pennsylvania State University, 110 Research Unit A, University Park, Pennsylvania 16802, USA
3. Department of Infectious Disease and Global Health, Cummings School of Veterinary Medicine, Tufts University, 200 Westboro Road, North Grafton, Massachusetts 01536, USA
4. Warnell School of Forestry and Natural Resources, University of Georgia, 180 E Green Street, Athens, Georgia 30602, USA