1. Ruminant Diseases and Immunology Research Unit, National Animal Disease Center, Agricultural Research Service, United States Department of Agriculture, 1920 Dayton Avenue, Ames, Iowa 50010, USA
2. Wyoming State Veterinary Laboratory, 1174 Snowy Range Road, Laramie, Wyoming 82070, USA
3. United States National Park Service, One Health Program, 1201 Oakridge Drive, Suite 200, Fort Collins, Colorado 80525, USA