1. Department of Pathobiology, University of Guelph, 491 Gordon Street, Guelph, Ontario N1G 2W1, Canada
2. Environmental and Life Sciences Graduate Program, Trent University, 1600 W Bank Drive, Peterborough,
3. Animal Health Laboratory, University of Guelph, 419 Gordon Street, Guelph, Ontario N1G 2W1, Canada