1. Saint Louis Zoo Institute for Conservation Medicine, One Government Drive, St. Louis, Missouri 63110, USA;
2. WildCare Institute, Saint Louis Zoo, One Government Drive, St. Louis, Missouri 63110, USA;
3. Max Planck Institute for Ornithology, Am Obstberg 1, D-78315, Radolfzell, Germany;
4. Whitney Harris World Ecology Center, University of Missouri–St. Louis, B216 Benton Hall, One University Boulevard, St. Louis, Missouri 63121, USA;
5. Department of Biology, Washington University in St. Louis, Campus Box 1137, One Brookings Drive, St. Louis, Missouri 63130, USA;
6. College of Veterinary Medicine, University of Florida, 2015 SW 16th Ave., Gainesville, Florida 32610, USA;