1. Wyoming Game and Fish Department, 528 S Adams, Laramie, Wyoming 82070, USA;
2. Colorado Division of Parks and Wildlife, Wildlife Health Program, 4330 Laporte Avenue, Fort Collins, Colorado 80521-2153, USA;
3. Wyoming Game and Fish Department, Wildlife Health Laboratory, 1174 Snowy Range Road, Laramie, Wyoming 82070, USA;
4. Wyoming Game and Fish Department, Thorne-Williams Wildlife Research Center, 2362 Hwy. 34, Wheatland, Wyoming 82201, USA;