1. Department of Ecology and Evolutionary Biology, University of Connecticut–Hartford Campus, 85 Lawler Road, West Hartford, Connecticut 06117, USA
2. Colorado State University, Western Slope Veterinary Diagnostic Laboratory, Grand Junction, Colorado 80154, USA
3. Colorado Division of Wildlife, Wildlife Research Center, Fort Collins, Colorado 80526, USA
4. Current address: Colorado State University Veterinary Diagnostic Laboratory, Fort Collins, Colorado, 80523-1644, USA
5. Colorado Division of Wildlife, 7405 Highway 50, Salida, Colorado 81201, USA
6. Biological Resource Management Division, National Park Service, Fort Collins, Colorado 80525, USA
7. Wind Cave National Park, Hot Springs, South Dakota 57747, USA
8. Parasitology, Colorado State University Veterinary Diagnostic Laboratory, Fort Collins, Colorado, 80523-1644