1. Department of Biological Sciences, Faculty of Science, Macquarie University, New South Wales 2109, Australia
2. Current address: Biosis Research Pty Ltd, 8 Tate St., Wollongong, New South Wales 2500, Australia
3. World Health Organization/Food and Agriculture Organization / Office International des Épizooties, Collaborating Centre for Reference and Research on Leptospirosis, Queensland Health Forensic and Scientific Services, Kessels Road, Coopers Plains, Queensland 4108, Australia