1. Department of Biology, University of Saskatchewan, 112 Science Place, Saskatoon, Saskatchewan S7N 5E2, Canada
2. Canadian Cooperative Wildlife Health Centre, Department of Veterinary Pathology, University of Saskatchewan, 52 Campus Drive, Saskatoon, Saskatchewan S7N 5B4, Canada
3. Prairie and Northern Wildlife Research Centre, Environment Canada, 115 Perimeter Road, Saskatoon, Saskatchewan S7N 0X4, Canada