1. Department of Veterinary Pathobiology, College of Veterinary Medicine, University of Illinois, Urbana, Illinois 61801, USA
2. San Diego Wild Animal Park, Escondido, California 92112, USA
3. Department of Land and Natural Resources, Division of Forestry and Wildlife, 2600 Piiholo Road, Makawao, Hawaii 96768, USA
4. Pacific Island Ecosystems Research Center, U. S. Geological Survey, Biological Resources Division, P.O. Box 218, Hawaii National Park, Hawaii 96718, USA