1. Department of Biology, University of Nebraska at Omaha, Omaha, Nebraska, 68182, USA
2. Current address: Wyoming Game and Fish Department, 117 S. Sublette Ave., Pinedale, Wyoming 82941, USA
3. Current address: Department of Natural Resource Ecology and Management, Iowa State University, Ames, Iowa 50011, USA
4. Utah Division of Wildlife Resources, 515 E 5300 S, Ogden, Utah 84405, USA
5. Current address: Minnesota Department of Natural Resources, 500 W. Lafayette Blvd., St. Paul, Minnesota 55155, USA