1. Department of Ecosystem and Public Health, Faculty of Veterinary Medicine, University of Calgary, 3280 Hospital Drive NW, Calgary, Alberta T2N4Z6, Canada
2. Department of Biological Sciences, University of Calgary, 2500 University Drive NW, Calgary, Alberta T2N1N4, Canada
3. Canadian Cooperative Wildlife Health Centre Alberta, 3280 Hospital Drive NW, Calgary, Alberta T2N4Z6, Canada
4. Ross University, P.O. Box 334, Basseterre, Saint Kitts, Saint Kitts and Nevis