1. Center for Conservation and Research of Endangered Wildlife, Cincinnati Zoo & Botanical Garden, Cincinnati, Ohio 45220, USA
2. Department of Fisheries and Wildlife, Michigan State University, East Lansing, Michigan 48824, USA
3. College of Veterinary Medicine and Biomedical Sciences, Colorado State University, Fort Collins, Colorado 80523, USA
4. Conservation and Research Center, National Zoological Park, Smithsonian Institution, Front Royal, Virginia 22630, USA
5. Mongolian Academy of Sciences, Ulaanbaatar, Mongolia