1. Department of Pathology, Microbiology & Immunology, School of Veterinary Medicine, University of California–Davis, Davis, California 95616, USA
2. 22835 Calle San Juan de Los Lagos, Moreno Valley, California 92553, USA
3. Department of Organismic Biology, Ecology, and Evolution, and Laboratory of Biomedical and Environmental Sciences, University of California–Los Angeles, Los Angeles, California 90024, USA
4. Current address: Department of Zoology, University of the Western Cape, Private Bag X17, Bellville, 7535, South Africa