1. National Wildlife Health Research Center, Hawaii Volcanoes Field Station, P.O. Box 218, Hawaii Volcanoes National Park, Hawaii 96718, USA
2. Department of Infectious Diseases, College of Veterinary Medicine, University of Florida, Gainesville, Florida 32610, USA
3. Department of Wildlife and Range Sciences, School of Forest Resources and Conservation, University of Florida, Gainesville, Florida 32610, USA
4. Florida Game and Fresh Water Fish Commission, 566 Commercial Boulevard, Naples, Florida 33942, USA