1. Florida Fish and Wildlife Cooperative Research Unit, Box 110485, Building 810, University of Florida, Gainesville, Florida 32611, USA
2. Current address: 17665 CR 833, Clewiston, Florida 33440, USA
3. Department of Infectious Diseases and Pathology, College of Veterinary Medicine, University of Florida, Box 110880, Gainesville, Florida 32610, USA
4. Department of Population Health, College of Veterinary Medicine, University of Georgia, 953 College Station Road, Athens, Georgia 30602, USA