1. Department of Biology, Stanford University, Stanford, California 94305, USA
2. Department of Otolaryngology, Head & Neck Surgery, Stanford University, Stanford, California 94305, USA
3. Department of Environmental Science, Policy and Management, University of California Berkeley, Berkeley, California 94720, USA
4. University of Namibia, Windhoek, Namibia
5. Oakland Zoo, Oakland, California 94605, USA
6. Department of Medicine, University of San Diego, San Diego, California 92093, USA
7. College of Veterinary Medicine, University of Tennessee, Knoxville, Tennessee 37919, USA
8. Department of Comparative Medicine, Stanford University, Stanford, California 94305, USA