1. Watershed Ecosystems Graduate Program, Trent University, Peterborough, Ontario K9J 7B8, Canada
2. Present Address: Institute of Zoology, Zoological Society of London. Regent's Park, London, NW1 4RY, UK
3. Canadian Wildlife Service, National Wildlife Research Centre, Carleton University, 1125 Colonel By Drive, Ottawa, Ontario K1A 0H3, Canada
4. Natural Resources DNA Profiling and Forensics Centre (NRDPFC), Trent University, Peterborough, Ontario K9J 7B8, Canada
5. Department of Biology, Trent University, Peterborough, Ontario K9J 7B8, Canada