1. School of Natural Sciences, Edith Cowan University, Joondalup, Perth 6027, Australia
2. Current address: CEH/SAFS, University Wales Bangor, Gwynedd LL57 2UP, UK
3. Department of Microbiology, QEII Medical Centre, The University of Western Australia, Nedlands, Perth 6009, Australia
4. Division of Veterinary and Biomedical Sciences, Murdoch University, Murdoch, Perth 6150, Australia