1. Department of Pathology, Ontario Veterinary College, University of Guelph, Guelph, Ontario, Canada N1G 2W1
2. Department of Environmental Biology, Ontario Agricultural College, University of Guelph, Guelph, Ontario, Canada N1G 2W1
3. Department of Population Medicine, Ontario Veterinary College, University of Guelph, Guelph, Ontario, Canada N1G 2W1