1. Institute of Arctic Biology, University of Alaska Fairbanks, Fairbanks, Alaska 99775-7000, USA
2. Current address: Dept. of Zoology and Physiology, University of Wyoming, Laramie, Wyoming 82071, USA.
3. Institute of Arctic Biology, and Department of Chemistry and Biochemistry, University of Alaska Fairbanks, Fairbanks, Alaska 99775-7000, USA
4. Institute of Arctic Biology, and Department of Biology and Wildlife, University of Alaska Fairbanks, Fairbanks, Alaska 99775-7000, USA