1. Colorado Division of Wildlife, Wildlife Research Center, 317 West Prospect Road, Fort Collins, Colorado 80526-2097, USA
2. Colorado Division of Wildlife, 300 West New York Avenue, Gunnison, Colorado 81230, USA
3. Colorado State University Diagnostic Laboratory, Department of Microbiology, Immunology, and Pathology, College of Veterinary Medicine, Colorado State University, Fort Collins, Colorado 80523, USA
4. Microbial Research, Incorporated, 2649 East Mulberry Street, Number 15, Fort Collins, Colorado 80524, USA