1. Ecological Modelling Services, P/L PO Box 6150, University of Queensland, St. Lucia, Queensland 4067, Australia
2. Pacific Islands Fisheries Science Center, NOAA Fisheries, 2570 Dole Street, Honolulu, Hawaii 96822, USA
3. US Geological Survey–National Wildlife Health Center, Honolulu Field Station, PO Box 50167, Honolulu, Hawaii 96850, USA