1. Washington Department of Fish and Wildlife, 2950 Ley Road, Cle Elum, Washington 98922, USA
2. Current address: US Geological Survey, New Mexico Cooperative Fisheries and Wildlife Research Unit, PO Box 30003 MSC 4901, Las Cruces, New Mexico 88003, USA
3. Washington Department of Fish and Wildlife, 16018 Mill Creek Boulevard, Mill Creek, Washington 98012, USA