1. Department of Ecology and Evolutionary Biology, University of Tennessee, 569 Dabney Hall, 1416 Circle Drive, Knoxville, Tennessee 37996, USA
2. Center for Microbial Genetics and Genomics, Northern Arizona University, 1298 S Knoles Drive, ARD Building 56, Room 213, Flagstaff, AZ 86011, USA
3. Current address: Department of Molecular, Cellular, and Biomedical Sciences, University of New Hampshire, Rudman Hall, 46 College Road, Durham, New Hampshire 03824, USA
4. Department of Forestry, Wildlife, and Fisheries, University of Tennessee, 274 Ellington Plant Science Building, Knoxville, Tennessee 37996, USA