1. University of British Columbia, School of Population and Public Health, 2206 E Mall, Vancouver, British Columbia V6T 1Z3, Canada
2. University of Glasgow, School of Medical, Veterinary, and Life Sciences, Wolfson Medical School Building, University of Glasgow, University Avenue, Glasgow, G12 8QQ, UK
3. Lilongwe Wildlife Trust, PO Box 1464, Lilongwe, Malawi
4. University of Southampton, School of Biological Sciences, Faculty of Environmental and Life Sciences, Life Sciences Building 85, University of Southampton, Highfield Campus, Southampton, SO17 1BJ, UK