1. Department of Forestry and Natural Resources, University of Kentucky, 105 T. P. Cooper Building, Lexington, Kentucky 40546, USA
2. Bear and Elk Program, Kentucky Department of Fish and Wildlife Resources, 1 Sportsman's Lane, Frankfort, Kentucky 40601, USA
3. Wildlife Health Program, Kentucky Department of Fish and Wildlife Resources, 1 Sportsman's Lane, Frankfort, Kentucky 40601, USA
4. Southeastern Cooperative Wildlife Disease Study and Department of Pathology, College of Veterinary Medicine, University of Georgia, 589 D. W. Brooks Drive, Athens, Georgia 30602, USA