1. Utah Division of Wildlife Resources, 1594 West North Temple, Suite 2110, Salt Lake City, Utah 84116, USA
2. Department of Wildlands Resources and Ecology Center, Utah State University, Logan, Utah 84322, USA
3. Utah Veterinary Diagnostic Laboratory, Department of Animal, Dairy, and Veterinary Sciences, Utah State University, 950 E 1400 N, Logan, Utah 84341, USA