1. Warnell School of Forestry and Natural Resources, The University of Georgia, 180 E Green St., Athens, Georgia 30602, USA
2. Southeastern Cooperative Wildlife Disease Study, Department of Population Health, College of Veterinary Medicine, The University of Georgia, 589 D. W. Brooks Dr., Athens, Georgia 30602, USA
3. Department of Biology and Institute for the Environment and Sustainability, The Miami University, 700 E High St., Oxford, Ohio 45056, USA
4. Department of Pathobiology, Ontario Veterinary College, University of Guelph, 50 Stone Rd. E, Guelph, Ontario N1G 2W1, Canada