1. Department of Pathobiology and Diagnostic Investigation, College of Veterinary Medicine, Michigan State University, Natural Resources Building Room 4, East Lansing, Michigan 48824, USA
2. US Geological Survey/Leetown Science Center, National Fish Health Research Laboratory, 11649 Leetown Road, Kearneysville, West Virginia 25430, USA
3. Department of Fisheries and Wildlife, College of Agriculture and Natural Resources, Michigan State University, 174 Food Safety and Toxicology Building, East Lansing, Michigan 48824, USA