1. Finnish Food Safety Authority Evira, Research and Laboratory Department, Production Animal and Wildlife Health Research Unit, Wildlife and Laboratory Section, Elektroniikkatie 5, 90590 Oulu, Finland;
2. Finnish Food Safety Authority Evira, Research and Laboratory Department, Veterinary Bacteriology Research Unit, Mustialankatu 3, 00790 Helsinki, Finland;
3. Natural Resources Institute Finland Luke, Itäinen Pitkäkatu 3, 20520 Turku, Finland;
4. Paris-Est University-ANSES, Animal Health Laboratory, Bacterial Zoonoses Unit, National Reference Centre for Human Brucellosis, National & EU/OIE/FAO Reference Laboratory for Animal Brucellosis, 14 rue Pierre et Marie Curie, 94706 Maisons-Alfort, France;