1. Southeastern Cooperative Wildlife Disease Study, 589 D. W. Brooks Drive, Wildlife Health Building, College of Veterinary Medicine, University of Georgia, Athens, Georgia 30602-4393, USA
2. Current address: Baker Institute for Animal Health, Department of Microbiology and Immunology, College of Veterinary Medicine, Cornell University, 235 Hungerford Hill Road, Ithaca, New York 14853, USA
3. Odum School of Ecology, University of Georgia, 140 E Green Street, Athens, Georgia 30602, USA
4. Retired