1. Veterinary Wildlife Services, South African National Parks, Kruger National Park, Private Bag X402, Skukuza 1350, South Africa
2. Department of Science and Innovation–National Research Foundation Centre of Excellence for Biomedical Tuberculosis Research, South African Medical Research Council Centre for TB Research, Division of Molecular Biology and Human Genetics, Faculty of M
3. Brain Function Research Group, School of Physiology, Faculty of Health Sciences, University of the Witwatersrand, 1 Jan Smuts Avenue, Johannesburg 2000, South Africa
4. Department of Clinical Sciences, College of Veterinary Medicine and Biomedical Science, Colorado State University, 1350 Centre Avenue, Fort Collins, Colorado 80521, USA
5. Department of Pathology and Laboratory Medicine, Schulich School of Medicine and Dentistry, Western University, 1151 Richmond Street, London, Ontario N6A 5C1, Canada;