1. Warnell School of Forestry and Natural Resources, The University of Georgia, 180 E Green Street, Athens, Georgia 30602, USA
2. Southeastern Cooperative Wildlife Disease Study, College of Veterinary Medicine, Department of Population Health, The University of Georgia, 589 D. W. Brooks Drive, Athens, Georgia 30602, USA
3. Florida Fish and Wildlife Conservation Commission, Fish and Wildlife Research Institute, 1105 SW Williston Road, Gainesville, Florida 32601, USA