1. Carlson College of Veterinary Medicine, Oregon State University, 700 SW 30th Street, Corvallis, Oregon 97331, USA
2. Department of Integrative Biology, Oregon State University, 2701 SW Campus Way, Corvallis, Oregon 97331, USA
3. Department of Veterinary Tropical Diseases, University of Pretoria, M35 Soutpan Road, Onderstepoord, Pretoria, 0110, South Africa
4. Department of Fisheries, Wildlife, and Conservation Sciences, 104 Nash Hall, 1500 SW Jefferson Avenue, Oregon State University, Corvallis, Oregon 97331, USA
5. College of Public Health, Medical and Veterinary Sciences, James Cook University, Townsville Queensland, Australia, 4814