1. Department of Wildland Resources, Utah State University, 5230 Old Main Hill, Logan, Utah 84322, USA
2. US Department of Agriculture, Wildlife Services, National Wildlife Research Center, 4101 LaPorte Avenue, Fort Collins, Colorado 80521, USA
3. Colorado Parks and Wildlife, Wildlife Health Research Center, 317 W Prospect Road, Fort Collins, Colorado 80525, USA
4. US Department of Agriculture, Wildlife Services, National Wildlife Research Center, Department of Wildland Resources, Utah State University, 5230 Old Main Hill, Logan, Utah 84322, USA