1. Colorado Division of Parks and Wildlife, Wildlife Health Program, 4330 Laporte Avenue, Fort Collins, Colorado 80521, USA
2. US Geological Survey, National Wildlife Health Center, 6060 Schroeder Road, Madison, Wisconsin 53706, USA
3. Department of Pathobiological Sciences, School of Veterinary Medicine, University of Wisconsin, 1655 Linden Dr., Madison, Wisconsin 53711, USA