1. Wildlife Management Branch, Environment and Sustainable Resource Development, 6909-116 St., Edmonton, Alberta T6H 4P2, Canada
2. Department of Biological Sciences, University of Alberta, Edmonton, Alberta T6G 2E9, Canada
3. Animal Health Branch, Agriculture and Rural Development, 6909-116 St., Edmonton, Alberta T6H 4P2, Canada
4. Animal Health Branch, Agriculture and Rural Development, 97 East Lake Ramp, Airdrie, Alberta T4A 0C3, Canada