1. School of Environment and Natural Resources, The Ohio State University, 2021 Coffey Rd., Columbus, Ohio 43210, USA
2. Smithsonian Tropical Research Institute, Apartado Postal 0843-03092 Panamá, República de Panamá
3. Ohio Division of Wildlife, 360 E. State Street, Athens, Ohio 45701, USA
4. Cornell University, Animal Health Diagnostic Center, 240 Farrier Rd., Ithaca, New York 14850, USA