1. Department of Biology, Tennessee Technological University, 1100 N Dixie Ave., Cookeville, Tennessee 38505, USA
2. Warnell School of Forestry and Natural Resources, University of Georgia, 180 E Green St., Athens, Georgia 30602, USA
3. Southeastern Cooperative Wildlife Disease Study, College of Veterinary Medicine, University of Georgia, 589 D. W. Brooks Dr., Athens, Georgia 30605, USA