1. Wildlife Health Center, University of California Davis, School of Veterinary Medicine, One Shields Avenue, Davis, California 95616, USA
2. Hubbs-SeaWorld Research Institute, 2595 Ingraham Street, San Diego, California 92109, USA
3. SeaWorld Parks and Entertainment, 500 SeaWorld Drive, San Diego, California 92109, USA
4. SeaWorld Parks and Entertainment, 7007 SeaWorld Drive, Orlando, Florida 32821, USA