1. Colorado Division of Parks and Wildlife, Wildlife Research Center, 317 West Prospect Road, Fort Collins, Colorado 80526-2097, USA
2. Newport Laboratories, 1525 Prairie Drive, Worthington, Minnesota 56187, USA
3. Wyoming Game and Fish Department, Wildlife Disease Laboratory, 1174, Snowy Range Road, Laramie, Wyoming 82070, USA
4. Department of Microbiology, Immunology, and Pathology, College of Veterinary Medicine and Biomedical Sciences, Colorado State University, Fort Collins, Colorado 80523-1619