1. Department of Pathobiology, University of Guelph, 50 Stone Road E, Guelph, Ontario N1G 2W1, Canada
2. Department of Population Medicine, University of Guelph, 50 Stone Road E, Guelph, Ontario N1G 2W1, Canada
3. Canadian Wildlife Health Cooperative, University of Guelph, 491 Gordon Street, Guelph, Ontario N1G 2W1, Canada
4. Southeastern Cooperative Wildlife Disease Study, College of Veterinary Medicine, University of Georgia, 589 D. W. Brooks Drive, Athens, Georgia 30602, USA