1. US Geological Survey, Alaska Science Center, 4210 University Dr., Anchorage, Alaska 99508, USA
2. Alaska Veterinary Pathology Services, 23834 The Clearing Drive, Eagle River, Alaska 99577, USA
3. US Geological Survey, National Wildlife Health Center, 6006 Schroeder Road, Madison, Wisconsin 53711, USA
4. Alaska Department of Fish and Game, 1300 College Road, Fairbanks, Alaska 99701, USA