1. Daniel B. Warnell School of Forestry and Natural Resources, University of Georgia, 180 E Green Street, , Athens, Georgia 30602, USA
2. Southeastern Cooperative Wildlife Disease Study, Department of Population Health, Wildlife Health Building, College of Veterinary Medicine, University of Georgia, 589 D. W. Brooks Drive, , Athens, Georgia 30602, USA
3. Joseph W. Jones Ecological Research Center, 3988 Jones Center Drive, Newton, Georgia 39870, USA
4. Current address: Georgia Department of Natural Resources, 1602 Lake Alexandria Lane, Thomasville, GA 31792, USA
5. Department of Biological Sciences, Auburn University, 331 Funchess Hall, Auburn, Alabama 36849, USA