1. School of Veterinary Medicine, Rakuno Gakuen University, 582 Bunkyodai Ebetsu, Hokkaido, 069-8501, Japan;
2. Department of Environmental and Symbiotic Science, Rakuno Gakuen University, 582 Bunkyodai Ebetsu, Hokkaido, 069-8501, Japan;
3. Hustai National Park, Mongolia, PO Box 1160, Ulaanbaatar 13, Mongolia;