1. University of Maryland Department of Biology; College Park Maryland 20742 USA
2. Michigan State University Department of Integrative Biology; Michigan State University Ecology, Evolutionary Biology and Behavior Program; East Lansing Michigan 48864 USA
3. Fisheries Resources Assessment and Monitoring Division; Northwest Fisheries Science Center; National Marine Fisheries Service; NOAA; Seattle Washington 98112 USA
4. Rutgers University Department of Marine and Coastal Sciences; New Brunswick New Jersey 08901 USA