1. School of Biological Sciences, University of Queensland, Brisbane 4072 Australia
2. Department of Fisheries and Wildlife, Oregon State University, 104 Nash Hall, Corvallis, Oregon 97331 USA
3. Department of Ecology and Evolution, Stony Brook University, Stony Brook, New York 11794-5245 USA
4. Department of Ecology, Evolution, and Marine Biology, University of California, Santa Barbara, California 93106 USA