1. School of Biological Sciences, University of Canterbury, Private Bag 4800, Christchurch 8140 New Zealand
2. School of Animal Biology, The University of Western Australia, 35 Stirling Highway, Crawley, Western Australia 6009 Australia
3. CSIRO Ecosystem Sciences, Centre for Environment and Life Sciences, Underwood Avenue, Floreat, Western Australia 6014 Australia
4. USDA-ARS Northern Plains Agricultural Research Laboratory, Sidney, Montana 59270 USA
5. Department of Life Sciences, Imperial College London, Silwood Park Campus, Buckhurst Road, Ascot, Berkshire SL5 7PY United Kingdom