1. Department of Fisheries and Wildlife; Oregon State University; Corvallis Oregon 97331 USA
2. Center for International Forestry Research (CIFOR); Jalan CIFOR; Situgede Bogor 16115 Indonesia
3. USDA Forest Service; Pacific Southwest Research Station; Institute of Pacific Islands Forestry; Hilo Hawaii 96720 USA
4. Department of Geophysics and Meteorology; Bogor Agricultural University; Darmaga Campus Bogor 16152 Indonesia
5. Soil Science Department; Sam Ratulangi University; Kampus Kleak-Bahu Manado 95115 Indonesia